Boechera paupercula |
Boechera divaricarpa |
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boechera paupercula, littleleaf rockcress, small-flower rock-cress, tiny suncress |
spreading-pod rock-cress, uplifting suncress |
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Habit | Perennials; long-lived; (cespitose); sexual; caudex often woody. | Biennials or perennials; short-lived; apomictic; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.3–1.5 dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, (0.07–)0.1–0.2 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)3–9 dm, glabrous or pubescent proximally, trichomes sessile, 2–4-rayed (some simple), to 0.7 mm, glabrous distally. |
Basal leaves | blade linear-oblanceolate, 1–3(–5) mm wide, margins entire, ciliate proximally, trichomes to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 2–6-rayed, (0.07–)0.1–0.2 mm. |
blade oblanceolate, 2–10 mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes sessile, 2–6-rayed, 0.1–0.4 mm. |
Cauline leaves | 2–6, rarely concealing stem; blade auricles 0 (or 0.2–1.5) mm, surfaces of distalmost leaves glabrous. |
(10–)15–56, concealing stem proximally; blade auricles 1–5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 3–8-flowered, unbranched. |
12–40(–65)-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous or sparsely pubescent; petals lavender to purplish, 4–6(–7) × 1–2 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals glabrous or with scattered trichomes; petals usually purple, rarely lavender, 6–9 × 1.5–3 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect, straight, 3–9 mm, glabrous or sparsely pubescent. |
divaricate-ascending to horizontal, straight, 5–12 mm, glabrous. |
Fruits | erect, appressed to rachis, not secund, straight, edges parallel, 2.5–5.5 cm × 1.3–1.7(–2) mm; valves glabrous; ovules 24–40 per ovary; style 0.2–1 mm. |
divaricate-ascending to horizontal, not appressed to rachis, not secund, straight, edges parallel, (4.5–)5.5–11 cm × 1.7–2.5 mm; valves glabrous; ovules 114–142 per ovary; style 0.05–0.2 mm. |
Seeds | usually uniseriate, rarely sub-biseriate, 1.5–2(–2.5) × 1–1.4 mm; wing continuous, 0.3–1 mm wide. |
uniseriate to sub-biseriate, 1.4–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 21. |
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Boechera paupercula |
Boechera divaricarpa |
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Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Rock outcrops, talus slopes, gravelly soil in alpine and subalpine habitats | Rock outcrops, talus slopes and gravelly hillsides in sagebrush, mountain shrub, and open conifer forests |
Elevation | 2400-3400 m (7900-11200 ft) | 900-2500 m (3000-8200 ft) |
Distribution |
CA; ID; OR; WA; WY
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CA; ID; MT; NV; UT; WA; WY
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Discussion | Boechera paupercula is usually subsumed under Arabis (Boechera) lyallii but is amply distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The name Arabis (Boechera) divaricarpa has been applied to nearly every hybrid containing a genome derived from B. stricta. This presents a serious barrier to understanding the evolution of Boechera and also is contrary to the International Code of Botanical Nomenclature, because some names usually placed in synonymy (i.e., B. grahamii and B. brachycarpa) have priority at species level (M. D. Windham and I. A. Al-Shehbaz 2007b). To address this problem, we treat the following as distinct species: B. acutina, B. grahamii (= B. brachycarpa of R. D. Dorn 2001), and B. pratincola (all considered synonyms of A. divaricarpa by R. C. Rollins 1993), and B. calderi, B. elkoensis, and B. quebecensis (taxa described after 1993). Detailed comparison among these taxa are provided by Windham and Al-Shehbaz (2007, 2007b). The narrow concept of B. divaricarpa advocated here encompasses apomictic triploid populations containing three distinct genomes, one each derived from B. retrofracta, B. sparsiflora, and B. stricta. If the species is defined more broadly, the name B. grahamii has priority. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 394. | FNA vol. 7, p. 374. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis paupercula, Arabis lyallii var. nubigena, Arabis microphylla var. nubigena, Arabis nubigena | Arabis divaricarpa, Arabis stokesiae |
Name authority | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 75. (2006) | (A. Nelson) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) |
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