Boechera oxylobula |
Boechera bodiensis |
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Bodie Hills rock cress |
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Habit | Perennials; short- to long-lived; (cespitose); sexual; caudex usually not woody. | Perennials; long-lived; (cespitose); apomictic; caudex somewhat woody. |
Stems | usually 3–7 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.4–2.5 dm, glabrous or pubescent proximally, trichomes simple and short-stalked, 2-rayed, 0.1–0.4 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–3.5 dm, densely pubescent proximally, trichomes short-stalked, 5–10-rayed, to 0.7 mm, sparsely pubescent or glabrous distally. |
Basal leaves | blade linear to linear-oblanceolate, 1–2.5 mm wide, margins usually entire, rarely denticulate, often ciliate, trichomes (simple), 0.3–0.7 mm, surfaces glabrous or sparsely pubescent, trichomes short-stalked, 2- or 3-rayed, 0.1–0.4 mm. |
blade narrowly oblanceolate, 1–3 mm wide, margins entire, ciliate proximally, trichomes to 0.7 mm, surfaces densely pubescent, trichomes short-stalked, 5–10-rayed, 0.08–0.25 mm. |
Cauline leaves | 3–12, not concealing stem; blade auricles absent, surfaces of distalmost leaves usually glabrous, rarely sparsely pubescent. |
4–9, not concealing stem; blade auricles 0.5–2 mm, surfaces of distalmost leaves pubescent. |
Racemes | 2–12-flowered, unbranched. |
8–25-flowered, usually unbranched. |
Flowers | ascending-divaricate at anthesis; sepals glabrous or pubescent; petals white to pale lavender, 4–5 × 1.5–2 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender to purple, 4–5 × ca. 1 mm, glabrous; pollen spheroid. |
Fruiting pedicels | divaricate-ascending to horizontal, slightly to strongly recurved, 3–8 mm, glabrous or sparsely pubescent, trichomes (isolated), simple. |
ascending to divaricate-ascending, straight, 3–7 mm, pubescent, trichomes appressed, branched. |
Fruits | pendent, not appressed to rachis, not or, rarely, weakly secund, straight, edges parallel, 1.5–3.5 cm × 1.2–2 mm; valves glabrous; ovules 28–44 per ovary; style 0.1–0.4 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight or curved, edges parallel, 4–6.2 cm × 1.2–1.8 mm; valves glabrous; ovules 48–68 per ovary; style 0.1–0.2 mm. |
Seeds | uniseriate, 0.9–1.2 × 0.6–1 mm; wing often continuous, 0.07–0.1 mm wide. |
uniseriate, 1–1.5 × 0.8–1.2 mm; wing continuous, 0.1–0.15 mm wide. |
Boechera oxylobula |
Boechera bodiensis |
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Phenology | Flowering May–Jul. | Flowering Jul–Aug. |
Habitat | Cliffs, rocky slopes, gravelly soil in sagebrush and open conifer forests | Loose soil, crevices of igneous rock |
Elevation | 2100-3600 m (6900-11800 ft) | 2400-2900 m (7900-9500 ft) |
Distribution |
CO |
CA; NV |
Discussion | Most of the collections assigned here have been called Arabis (Boechera) demissa by other authors (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b). Because the holotype of A. demissa is identical to B. oxylobula in nearly every way (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison), we treat them as conspecific. Boechera oxylobula is restricted to Garfield, Gunnison, Hinsdale, Lake, Mineral, Park, and Saguache counties in central Colorado. The taxon traditionally treated as A. (Boechera) demissa var. languida is here recognized as an apomictic species of hybrid origin (see Windham and Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. In his original description, Rollins suggested that Boechera bodiensis is closely related to B. cobrensis and B. falcifructa. Subsequent studies (M. D. Windham and I. A. Al-Shehbaz, unpubl.) indicate that both B. bodiensis and B. falcifructa are apomictic hybrids containing one or more genomes derived from B. cobrensis. In the case of B. falcifructa, the second parent is clearly B. fernaldiana, but the second parent of B. bodiensis remains uncertain. Boechera bodiensis is most similar to B. falcifructa. The latter differs by having gently recurved fruiting pedicels 6–12 mm, 62–80 ovules per ovary, and pendulous fruits; it is known from Elko and Lander counties, Nevada. By contrast, B. bodiensis has straight fruiting pedicels 3–7 mm long, divaricate-ascending fruits, 48–68 ovules per ovary, and is known from Mono County, California, and Lyon and Mineral counties, Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 391. | FNA vol. 7, p. 367. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis oxylobula, Arabis aprica, Arabis demissa, Arabis rugocarpa, B. demissa | Arabis bodiensis |
Name authority | (Greene) W. A. Weber: Phytologia 51: 370. (1982) | (Rollins) Al-Shehbaz: Novon 13: 384. (2003) |
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