Boechera lemmonii |
Boechera macounii |
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Lemmon's rock-cress, Lemmon's suncress, wind-blown suncress |
littleleaf suncress, Macoun's suncress, sloppy suncress |
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Habit | Perennials; long-lived; (somewhat cespitose); sexual or apomictic; caudex woody. | Perennials; usually long-lived; apomictic; caudex often woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.5–2(–2.5) dm, glabrous or sparsely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, glabrous distally. |
1 or several per caudex branch, arising from center or margin of rosette near ground surface, 2–3.6 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, 0.1–0.8 mm, glabrous distally. |
Basal leaves | blade oblanceolate to obovate, 1.5–5 mm wide, margins usually entire, rarely slightly dentate, ciliate along petiole, surfaces densely to sparsely pubescent, trichomes short-stalked, 3–9-rayed, 0.1–0.2 mm. |
blade oblanceolate to linear-oblanceolate, 1–3(–5) mm wide, margins entire or minutely toothed, ciliate proximally, trichomes (simple), to 0.7 mm, surfaces densely pubescent, trichomes short-stalked, 3–6-rayed, 0.07–0.3 mm. |
Cauline leaves | 2–8(–12), not concealing stem; blade auricles absent or 0.1–0.5 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
(5–)9–16, often concealing stem proximally; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 3–12(–17)-flowered, usually unbranched. |
10–33-flowered, usually unbranched. |
Flowers | divaricate-ascending at anthesis; sepals glabrous or sparsely pubescent; petals purple to lavender, 3.5–6 × 1–1.5 mm; pollen ellipsoid or spheroid. |
divaricate-ascending at anthesis; sepals pubescent; petals lavender to purple, 4–5 × 0.5–0.8 mm, glabrous; pollen spheroid. |
Fruiting pedicels | divaricate-ascending to slightly descending, usually slightly recurved, rarely straight, 2–6 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
divaricate-ascending to horizontal, straight, 5–13(–17) mm, usually densely pubescent, rarely glabrous, trichomes appressed, 2- or 3-rayed. |
Fruits | divaricate-ascending to slightly descending, not appressed to rachis, secund, straight or curved, edges parallel, (1.6–) 2–4.4 cm × 1.6–2.3 mm; valves glabrous; ovules 28 40(–44) per ovary; style 0.1–0.2 mm. |
divaricate-ascending to descending, not appressed to rachis, not secund, usually curved, edges parallel, 3.5–6.5 cm × 0.8–1.2 mm; valves glabrous; ovules 56–78 per ovary; style 0.1–0.4 mm. |
Seeds | uniseriate, 1.3–2 × 1–1.5 mm; wing continuous, 0.1–0.5 mm wide. |
uniseriate, 1–1.2 × 0.7–1 mm; wing often continuous, 0.05–0.15 mm wide. |
2n | = 14. |
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Boechera lemmonii |
Boechera macounii |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Cliffs, talus slopes, and gravelly soil in alpine and subalpine habitats | Rocky hillsides in open pinewoods |
Elevation | 2100-4400 m (6900-14400 ft) | 2200-2900 m (7200-9500 ft) |
Distribution |
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; YT
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ID; MT; WY; BC |
Discussion | Boechera lemmonii is easily recognized by its combination of secund fruits, mat-forming habit, purplish sepals, and obovate-oblanceolate cauline leaves. Both sexual and apomictic collections are known; further study is needed to determine whether they truly are conspecific. The taxa traditionally treated as Arabis (Boechera) lemmonii vars. depauperata, drepanoloba, and paddoensis are apomictic hybrids here recognized as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera macounii is an apomictic species that arose through hybridization between B. collinsii and B. microphylla (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 388. | FNA vol. 7, p. 389. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis lemmonii, Arabis bracteolata, Arabis canescens var. latifolia, Arabis codyi, Arabis egglestonii, Arabis kennedyi, Arabis latifolia, Arabis oreocallis, Arabis polyclada, Arabis semisepulta | Arabis macounii, Arabis densicaulis, Arabis microphylla var. macounii |
Name authority | (S. Watson) W. A. Weber: Phytologia 51: 370. (1982) | (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 267. (2007) |
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