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Brewer's rock-cress

Habit Perennials; long-lived; sexual; caudex often woody. Perennials; long-lived; sexual; caudex woody.
Stems

1 or few per caudex branch, arising from center or margin of rosette, somewhat elevated on woody base or near ground surface, (0.7–)2–3.5 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.05–0.15 mm, moderately to sparsely pubescent distally.

usually 1 per caudex branch, arising from center of rosette, somewhat elevated on woody base or from ground surface, 0.6–3.5(–4.5) dm, densely pubescent proximally, trichomes long-stalked and simple, 2-rayed (sometimes all simple), 0.4–1 mm, sparsely pubescent or glabrous distally.

Basal leaves

blade narrowly oblanceolate, 2–5 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.05–0.15 mm.

blade oblanceolate, 3–7(–11) mm wide, margins entire or dentate, ciliate proximally, trichomes to 1 mm, surfaces densely pubescent, trichomes long-stalked, 2–4-rayed, 0.4–0.8 mm.

Cauline leaves

4–10, often concealing stem proximally;

blade auricles absent, surfaces of distalmost leaves sparsely to moderately pubescent.

5–15(–28), often concealing stem proximally;

blade auricles 0.5–3(–5) mm, surfaces of distalmost leaves pubescent.

Racemes

5–12-flowered, usually unbranched.

7–20(–30)-flowered, usually unbranched.

Flowers

divaricate at anthesis;

sepals pubescent;

petals purple to lavender, 6–10 × 1.5–2.5 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals usually purple (rarely lavender), 7–12 × 2–4 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

descending, recurved, 4–14 mm, pubescent, trichomes appressed, branched.

ascending to divaricate-ascending, straight, 3–25 mm, pubescent or glabrous, trichomes spreading, simple and 2-rayed.

Fruits

pendent, usually not appressed to rachis, rarely secund, straight or curved, 2–5 cm × 1.7–2.2 mm;

valves glabrous or pubescent throughout;

ovules 26–36 per ovary;

style 0.2–0.9 mm.

ascending to divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.5–10 cm × 1.5–2.2 mm;

valves usually glabrous, rarely sparsely pubescent;

ovules 48–96 per ovary;

style 0.05–0.3 mm.

Seeds

uniseriate, 1.5–2 × 1–1.5 mm;

wing distal or continuous, 0.1–0.2 mm wide.

uniseriate, 1.2–1.7 × 1–1.2 mm;

wing continuous, 0.1–0.2 mm wide distally.

Boechera lasiocarpa

Boechera breweri

Phenology Flowering May–Jun.
Habitat Rocky ridges and slopes with dwarf sagebrush
Elevation 1800-2800 m (5900-9200 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera lasiocarpa is a sexual species that appears to be restricted to the Bear River Mountains and northern Wasatch Range in north-central Utah. Reports of Arabis (Boechera) lasiocarpa from central Idaho (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b) represent B. rollinsiorum (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Relatively few other sexual diploid species of Boechera co-occur with B. breweri, and the only currently recognized hybrid involving this taxon is B. rubicundula (= B. arcuata × B. breweri; M. D. Windham and I. A. Al-Shehbaz 2007). Boechera breweri appears to be related to B. sparsiflora, and some specimens of subsp. shastaensis can be difficult to separate from that species. The two subspecies appear to intergrade on the western side of the Sacramento Valley from Tehama to Solano counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems 0.6-2 dm, pubescent distally; fruiting pedicels 3-8(-12) mm, pubescent; fruits 3.5-7.5 cm.
subsp. breweri
1. Stems 1.8-3.5(-4.5) dm, glabrous distally; fruiting pedicels (7-)10-25 mm, usually glabrous; fruits (4.5-)7-10 cm.
subsp. shastaensis
Source FNA vol. 7, p. 387. FNA vol. 7, p. 367.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Subordinate taxa
B. breweri subsp. breweri, B. breweri subsp. shastaensis
Synonyms Arabis lasiocarpa Arabis breweri, Arabis breweri var. figularis, Arabis epilobioides
Name authority (Rollins) Dorn: Brittonia 55: 3. (2003) (S. Watson) Al-Shehbaz: Novon 13: 384. (2003)
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