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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress

bristly-leaf rock-cress

Habit Perennials; long-lived; sexual; caudex usually woody. Biennials or perennials; short-lived; sexual; caudex present or absent.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout.

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–8(–10) dm, usually glabrous proximally, rarely sparsely pubescent, trichomes simple, to 0.7 mm, glabrous distally.

Basal leaves

blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous.

blade oblanceolate, 3–8(–13) mm wide, margins entire or denticulate, ciliate, trichomes (simple), 0.5–1.2 mm, surfaces sparsely to densely pubescent, trichomes simple, to 0.8 mm, often mixed with short-stalked, 2- or 3-rayed ones, 0.15–0.5 mm.

Cauline leaves

2–4, not concealing stem;

blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally.

6–30(–45), often concealing stem proximally;

blade auricles 1–1.5 mm, surfaces of distalmost leaves sparsely pubescent, sometimes margins ciliate.

Racemes

2–5-flowered, unbranched.

10–30(–50)-flowered, usually unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous;

petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous;

pollen ellipsoid.

pendent at anthesis;

sepals pubescent;

petals white, 3–4 × 0.7–1.2 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

ascending, straight, 4–10 mm, glabrous.

reflexed, abruptly recurved at base, otherwise straight, 4–10 mm, glabrous.

Fruits

suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm;

valves glabrous;

ovules 10–20 per ovary;

style 0.05–0.3 mm.

strongly reflexed, usually appressed to rachis, not secund, straight, edges parallel, 5–9 cm × 1.8–2.5 mm;

valves glabrous;

ovules 46–80 per ovary;

style 0.2–0.8 mm.

Seeds

uniseriate, 3–6 × 2–4 mm;

wing continuous, 1.3–2.5 mm wide.

uniseriate, 1.8–2.1 × 1–1.5 mm;

wing continuous, 0.15–0.25 mm wide.

Boechera howellii

Boechera rectissima

Phenology Flowering Jun–Aug. Flowering May–Jul.
Habitat Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats Rocky slopes in open conifer forests
Elevation 1800-3800 m (5900-12500 ft) 1500-2500 m (4900-8200 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera rectissima is easily recognized by its combination of glabrous stems, basal leaves with large, predominantly simple trichomes, short (3–4 mm), white petals, and strongly reflexed, appressed fruits. Plants assigned to var. simulans appear to be hybrids between this species and B. retrofracta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 384. FNA vol. 7, p. 401.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata Arabis rectissima, Arabis setigera, Arabis wyndii
Name authority (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) (Greene) Al-Shehbaz: Novon 13: 388. (2003)
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