Boechera howellii |
Boechera puberula |
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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress |
Blue Mountain rockcress, hoary rock-cress, silver rockcress |
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Habit | Perennials; long-lived; sexual; caudex usually woody. | Perennials; short-lived; sexual; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3(–0.5) mm, similarly pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous. |
blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm. |
Cauline leaves | 2–4, not concealing stem; blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally. |
7–45(–65), concealing stem proximally; blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent. |
Racemes | 2–5-flowered, unbranched. |
10–40(–64)-flowered, usually unbranched. |
Flowers | erect to ascending at anthesis; sepals glabrous; petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous; pollen ellipsoid. |
ascending to descending at anthesis; sepals pubescent; petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | ascending, straight, 4–10 mm, glabrous. |
pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched. |
Fruits | suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm; valves glabrous; ovules 10–20 per ovary; style 0.05–0.3 mm. |
closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm; valves pubescent throughout; ovules 38–64 per ovary; style 0.05–0.1 mm. |
Seeds | uniseriate, 3–6 × 2–4 mm; wing continuous, 1.3–2.5 mm wide. |
uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.3 mm wide. |
2n | = 14. |
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Boechera howellii |
Boechera puberula |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Jul. |
Habitat | Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats | Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities |
Elevation | 1800-3800 m (5900-12500 ft) | 1300-2900 m (4300-9500 ft) |
Distribution |
CA; NV; OR
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CA; NV; OR; UT
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Discussion | Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 384. | FNA vol. 7, p. 399. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata | Arabis puberula, Arabis arida, Arabis beckwithii, Arabis lignipes var. impar, Arabis sabulosa, Arabis subpinnatifida var. beckwithii, Arabis subpinnatifida var. impar, B. beckwithii |
Name authority | (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) | (Nuttall) Dorn: Brittonia, 55: 3. (2003) |
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