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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress

meadow rock-cress

Habit Perennials; long-lived; sexual; caudex usually woody. Perennials; usually short-lived; apomictic; caudex not woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout.

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6 dm, pubescent proximally, trichomes sessile, 2- or 3-rayed, 0.15–0.6 mm, glabrous distally.

Basal leaves

blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous.

blade oblanceolate, 1.5–5(–7) mm wide, margins usually entire, rarely denticulate, ciliate at petiole base, trichomes to 0.4 mm, surfaces pubescent, trichomes sessile, 2–4(–6)-rayed, 0.1–0.45 mm.

Cauline leaves

2–4, not concealing stem;

blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally.

(7–)13–42, concealing stem proximally;

blade auricles 1–4 mm, surfaces of distalmost leaves sparsely pubescent apically.

Racemes

2–5-flowered, unbranched.

(7–)14–45-flowered, usually unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous;

petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals glabrous;

petals lavender, 6–10 × 1.5–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

ascending, straight, 4–10 mm, glabrous.

erect, straight, 5–12 mm, glabrous.

Fruits

suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm;

valves glabrous;

ovules 10–20 per ovary;

style 0.05–0.3 mm.

erect, appressed to rachis, not secund, straight, edges parallel, 4–6.5 cm × 1.5–2 mm;

valves glabrous;

ovules 60–140 per ovary;

style 0.1–0.3 mm.

Seeds

uniseriate, 3–6 × 2–4 mm;

wing continuous, 1.3–2.5 mm wide.

sub-biseriate, 1.4–2 × 1–1.4 mm;

wing continuous, 0.1–0.35 mm wide.

Boechera howellii

Boechera pratincola

Phenology Flowering Jun–Aug. Flowering Jun–Aug.
Habitat Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats Rocky slopes and soil patches in open areas and along forest edges
Elevation 1800-3800 m (5900-12500 ft) 1900-3200 m (6200-10500 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera pratincola is an apomictic species that arose through hybridization between B. paupercula and B. stricta (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 384. FNA vol. 7, p. 399.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata Arabis pratincola, Arabis drummondii var. pratincola, Arabis nemophila
Name authority (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 245. (2007)
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