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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress

nodding rock cress, rabbit-ear rockcress

Habit Perennials; long-lived; sexual; caudex usually woody. Perennials; short- to long-lived; sexual or apomictic; caudex sometimes woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout.

usually 2–6 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.6–3(–3.7) dm, sparsely to densely pubescent proximally, trichomes simple, 0.3–0.8 mm, glabrous distally.

Basal leaves

blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous.

blade oblanceolate to obovate, 1.5–6 mm wide, margins usually entire, rarely dentate, ciliate throughout, trichomes (usually simple), 0.4–1 mm, surfaces pubescent, trichomes simple and short- and long-stalked, 2-rayed, 0.3–0.8 mm.

Cauline leaves

2–4, not concealing stem;

blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally.

2–10(–13), not concealing stem;

blade auricles absent or, rarely, to 0.7 mm, surfaces of distalmost leaves glabrous or margins sparsely ciliate.

Racemes

2–5-flowered, unbranched.

4–14-flowered, usually unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous;

petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous;

pollen ellipsoid.

divaricate-ascending at anthesis;

sepals usually sparsely pubescent, rarely glabrous;

petals whitish to pale lavender, 4–6 × 1–1.5 mm, glabrous;

pollen ellipsoid or spheroid.

Fruiting pedicels

ascending, straight, 4–10 mm, glabrous.

divaricate-ascending to horizontal, curved or angled downward, 3–7(–10) mm, usually glabrous, rarely with some simple trichomes.

Fruits

suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm;

valves glabrous;

ovules 10–20 per ovary;

style 0.05–0.3 mm.

widely pendent, not appressed to rachis, not secund, curved to nearly straight, edges parallel, 2.2–4 cm × 1.2–2.1 mm;

valves glabrous;

ovules 40–70(–90) per ovary;

style 0.1–0.3(–0.5) mm.

Seeds

uniseriate, 3–6 × 2–4 mm;

wing continuous, 1.3–2.5 mm wide.

biseriate, 0.9–1.2 × 0.6–0.9 mm, usually not winged.

2n

= 14, 21.

Boechera howellii

Boechera pendulina

Phenology Flowering Jun–Aug. Flowering Apr–Jun.
Habitat Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats Rock outcrops, open gravelly flats and hillsides in sagebrush, pinyon-juniper, mountain mahogany, open conifer forests
Elevation 1800-3800 m (5900-12500 ft) 1600-3100 m (5200-10200 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; NV; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although included in Arabis (Boechera) demissa by some authors (e.g., S. L. Welsh et al. 2003; N. H. Holmgren 2005b), B. pendulina is readily distinguished from that species (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). Typical collections are sexual diploids, whereas the type of var. russeola is a triploid apomict; further study is needed to determine if the two are conspecific.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 384. FNA vol. 7, p. 395.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata Arabis pendulina, Arabis demissa var. russeola, Arabis diehlii, Arabis pendulina var. russeola, Arabis setulosa, B. demissa var. pendulina, B. demissa var. russeola, B. pendulina var. russeola
Name authority (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) (Greene) W. A. Weber: Phytologia 51: 370. (1982)
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