Boechera howellii |
Boechera languida |
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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress |
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Habit | Perennials; long-lived; sexual; caudex usually woody. | Perennials; short- to long-lived; apomictic; caudex usually not, or, rarely, woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout. |
usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 1–3 dm, pubescent proximally, trichomes simple, to 1 mm, mixed with short-stalked, 2-rayed ones, 0.15–0.5 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous. |
blade linear-oblanceolate to oblanceolate, 1–5(–9) mm wide, margins entire, ciliate, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes simple and 2–4-rayed, 0.15–0.5 mm. |
Cauline leaves | 2–4, not concealing stem; blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally. |
3–7, not concealing stem; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves sparsely pubescent or glabrous. |
Racemes | 2–5-flowered, unbranched. |
10–20-flowered, usually unbranched. |
Flowers | erect to ascending at anthesis; sepals glabrous; petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals white to pale lavender, 4–6 × 1.5–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | ascending, straight, 4–10 mm, glabrous. |
horizontal, slightly to strongly recurved near apex, 3–13 mm, glabrous or, rarely, with simple trichomes. |
Fruits | suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm; valves glabrous; ovules 10–20 per ovary; style 0.05–0.3 mm. |
pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, 3–4.5 cm × 1.8–2 mm; valves glabrous; ovules 42–70 per ovary; style 0.1–0.3 mm. |
Seeds | uniseriate, 3–6 × 2–4 mm; wing continuous, 1.3–2.5 mm wide. |
sub-biseriate, 1.2–1.5 × 0.8–1 mm; wing absent or distal, 0.05–0.12 mm wide. |
Boechera howellii |
Boechera languida |
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Phenology | Flowering Jun–Aug. | Flowering May–Jun. |
Habitat | Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats | Rocky slopes in sagebrush and pinyon-juniper communities |
Elevation | 1800-3800 m (5900-12500 ft) | 1800-2300 m (5900-7500 ft) |
Distribution |
CA; NV; OR
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WY |
Discussion | Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera languida is an apomictic hybrid that is often subsumed under Arabis demissa (= B. oxylobula). Morphological evidence suggests a closer relationship to B. spatifolia, which probably is one of its parents (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 384. | FNA vol. 7, p. 387. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata | Arabis demissa var. languida, B. demissa var. languida |
Name authority | (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) | (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 244. (2007) |
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