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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress

Habit Perennials; long-lived; sexual; caudex usually woody. Biennials or perennials; short-lived; sexual; caudex present or absent.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout.

usually 1 per caudex branch, arising from center of rosette near ground surface, rarely arising laterally proximal to sterile shoots, 2.5–8.5 dm, densely pubescent proximally, trichomes simple or spurred, 0.9–1.5 mm, glabrous distally.

Basal leaves

blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous.

blade oblanceolate, 5–12 mm wide, margins shallowly dentate, sometimes ciliate near petiole base, surfaces sparsely to densely pubescent, trichomes short- to long-stalked, 2- or 3-rayed, usually 0.3–0.6 mm.

Cauline leaves

2–4, not concealing stem;

blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally.

30–65, often concealing stem proximally;

blade auricles 0.8–3 mm, surfaces of distalmost leaves usually glabrous.

Racemes

2–5-flowered, unbranched.

(12–)15–50-flowered, usually unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous;

petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals white to pale lavender, 6–9 × 0.6–1 mm;

valves glabrous;

pollen ellipsoid.

Fruiting pedicels

ascending, straight, 4–10 mm, glabrous.

divaricate-ascending or horizontal, recurved, (15–)20–47 mm, glabrous.

Fruits

suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm;

valves glabrous;

ovules 10–20 per ovary;

style 0.05–0.3 mm.

widely pendent, not appressed to rachis, not secund, curved, edges parallel, (2.5–)3–7.5 cm × 1.5–2.8 mm, glabrous;

ovules 130–210 per ovary;

style 0.2–0.5 mm.

Seeds

uniseriate, 3–6 × 2–4 mm;

wing continuous, 1.3–2.5 mm wide.

biseriate, 1.2–1.4 × 0.7–0.9 mm;

wing continuous or distal, to 0.2 mm wide.

Boechera howellii

Boechera gracilipes

Phenology Flowering Jun–Aug. Flowering Apr–Jun.
Habitat Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats Basalt, limestone, and sandy soils in ponderosa pine forests and pinyon-juniper woodlands
Elevation 1800-3800 m (5900-12500 ft) 1700-2300 m (5600-7500 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NV; UT
[BONAP county map]
Discussion

Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fruiting pedicel length, considered diagnostic by R. C. Rollins (1993) and N. H. Holmgren (2005b), occasionally fails to separate Boechera gracilipes from the closely-related B. fendleri. The two are consistently distinguished by trichome characters. In B. gracilipes, basal leaves usually lack prominent cilia and surfaces are persistently pubescent with at least some 3-rayed trichomes. Also, stems are rather densely pilose proximally, with the largest trichomes more than 0.9 mm. By contrast, basal leaves of B. fendleri always have prominent cilia, surfaces are often glabrescent, and 3-rayed trichomes usually are rare or absent. Stems of the latter are proximally hirsute to hispid, with the largest trichomes less than 0.9 mm.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 384. FNA vol. 7, p. 381.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata Arabis gracilipes, Arabis arcuata var. longipes, Arabis perennans var. longipes
Name authority (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) (Greene) Dorn: Brittonia 55: 3. (2003)
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