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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress

Habit Perennials; long-lived; sexual; caudex usually woody. Perennials; usually long-lived; sexual; caudex often woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout.

1 to several per caudex branch, arising from center and margin of rosette, slightly elevated on woody base or from near ground surface, 0.8–3 dm, densely pubescent throughout, trichomes short-stalked, 5–10-rayed, 0.1–0.2 mm.

Basal leaves

blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous.

blade linear-oblanceolate, 1–5 mm wide, margins entire or with few teeth subapically, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–10-rayed, 0.1–0.2 mm.

Cauline leaves

2–4, not concealing stem;

blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally.

5–18, concealing stem proximally;

blade auricles 0.5–2.5 mm, surfaces of distalmost leaves densely pubescent.

Racemes

2–5-flowered, unbranched.

8–35-flowered, usually unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous;

petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous;

pollen ellipsoid.

erect to ascending at anthesis;

sepals pubescent;

petals purplish, 8–12 × 2.5–4.5 mm;

pollen ellipsoid.

Fruiting pedicels

ascending, straight, 4–10 mm, glabrous.

erect to ascending, straight, 5–15 mm, pubescent, trichomes appressed, branched.

Fruits

suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm;

valves glabrous;

ovules 10–20 per ovary;

style 0.05–0.3 mm.

erect to ascending, often appressed to rachis, not secund, straight, edges parallel, 3–5.8 cm × 1.4–1.8 mm;

valves densely pubescent throughout;

ovules 34–68 per ovary;

style 1–1.5 mm.

Seeds

uniseriate, 3–6 × 2–4 mm;

wing continuous, 1.3–2.5 mm wide.

uniseriate, 1–1.5 × 0.8–1.2 mm;

wing continuous, 0.5–0.1 mm wide.

Boechera howellii

Boechera fecunda

Phenology Flowering Jun–Aug. Flowering Apr–Jun.
Habitat Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats Rocky ground in sagebrush areas
Elevation 1800-3800 m (5900-12500 ft) 1200-1800 m (3900-5900 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
MT
[BONAP county map]
Discussion

Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Boechera fecunda is a distinctive sexual species known from Ravali and Silver Bow counties.

Boechera fecunda is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 384. FNA vol. 7, p. 377.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata Arabis fecunda
Name authority (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) (Rollins) Dorn: Brittonia 55: 3. (2003)
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