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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress

hidden rockcress

Habit Perennials; long-lived; sexual; caudex usually woody. Perennials; long-lived; (cespitose); sexual; caudex woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout.

usually 1 per caudex branch, arising from center of rosette near ground surface, 1–2.5 dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.2–0.6 mm, and sparse, simple ones to 1 mm, glabrous distally.

Basal leaves

blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous.

blade narrowly oblanceolate, 1.5–3 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.2 mm.

Cauline leaves

2–4, not concealing stem;

blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally.

3–5, not concealing stem;

blade auricles 0.5–1 mm, surfaces of distalmost leaves pubescent.

Racemes

2–5-flowered, unbranched.

10–22-flowered, unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous;

petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals white, 3–4 × 0.5–0.8 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

ascending, straight, 4–10 mm, glabrous.

divaricate-ascending, straight, 4–8 mm, sparsely pubescent, trichomes appressed, branched.

Fruits

suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm;

valves glabrous;

ovules 10–20 per ovary;

style 0.05–0.3 mm.

divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 3–4 cm × 1.2–1.5 mm;

valves glabrous;

ovules 46–52 per ovary;

style ca. 0.1 mm.

Seeds

uniseriate, 3–6 × 2–4 mm;

wing continuous, 1.3–2.5 mm wide.

uniseriate, 1–1.1 × 0.9–1 mm;

wing continuous, ca. 0.1 mm wide.

Boechera howellii

Boechera evadens

Phenology Flowering Jun–Aug. Flowering Jun.
Habitat Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats Rock outcrops
Elevation 1800-3800 m (5900-12500 ft) ca. 2600 m (ca. 8500 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera evadens is a sexual taxon that was originally interpreted as a disjunct population of Arabis (Boechera) fernaldiana, but is easily distinguished from that species by having much shorter (3–4 versus 7–12 mm) petals, larger (0.2–0.6 versus 0.04–0.1 mm) trichomes proximally on stems, and shorter (3–4 versus 4–7.5 cm) fruits (see M. D. Windham and I. A. Al-Shehbaz 2006 for additional distinctions). It is known only from the holotype specimen collected near Sherman Pass in Tulare County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 384. FNA vol. 7, p. 376.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata
Name authority (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 66. (2006)
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