Boechera crandallii |
Boechera gracilipes |
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Crandall's rockcress |
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Habit | Perennials; long-lived; (often cespitose); sexual; caudex somewhat woody. | Biennials or perennials; short-lived; sexual; caudex present or absent. |
Stems | usually 2–5 per caudex branch, arising from margin of rosette near ground surface, (1–)1.5–4 dm, densely pubescent proximally, trichomes short-stalked, 5–8-rayed, 0.1–0.2 mm, moderately to sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, rarely arising laterally proximal to sterile shoots, 2.5–8.5 dm, densely pubescent proximally, trichomes simple or spurred, 0.9–1.5 mm, glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1.5–3(–5) mm wide, margins entire, ciliate along petiole base, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 5–8-rayed 0.1–0.2 mm. |
blade oblanceolate, 5–12 mm wide, margins shallowly dentate, sometimes ciliate near petiole base, surfaces sparsely to densely pubescent, trichomes short- to long-stalked, 2- or 3-rayed, usually 0.3–0.6 mm. |
Cauline leaves | 5–14, not concealing stem; blade auricles 0.1–0.5 mm, surfaces of distalmost leaves pubescent. |
30–65, often concealing stem proximally; blade auricles 0.8–3 mm, surfaces of distalmost leaves usually glabrous. |
Racemes | 10–30-flowered, usually unbranched. |
(12–)15–50-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals usually white, 5–7 × 1–2 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white to pale lavender, 6–9 × 0.6–1 mm; valves glabrous; pollen ellipsoid. |
Fruiting pedicels | ascending to divaricate-ascending, straight, 5–10 mm, pubescent, trichomes branched. |
divaricate-ascending or horizontal, recurved, (15–)20–47 mm, glabrous. |
Fruits | ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 3–5.5 cm × 0.9–1.2 mm; valves glabrous; ovules 56–84 per ovary; style 0.1–0.5 mm. |
widely pendent, not appressed to rachis, not secund, curved, edges parallel, (2.5–)3–7.5 cm × 1.5–2.8 mm, glabrous; ovules 130–210 per ovary; style 0.2–0.5 mm. |
Seeds | uniseriate, 0.8–1 × 0.6–0.9 mm; wing continuous (rarely absent), to 0.1 mm wide. |
biseriate, 1.2–1.4 × 0.7–0.9 mm; wing continuous or distal, to 0.2 mm wide. |
2n | = 14. |
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Boechera crandallii |
Boechera gracilipes |
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Phenology | Flowering May–Jun. | Flowering Apr–Jun. |
Habitat | Rocky slopes and gravelly soil in sagebrush, mountain shrub, open conifer forests | Basalt, limestone, and sandy soils in ponderosa pine forests and pinyon-juniper woodlands |
Elevation | 2000-2700 m (6600-8900 ft) | 1700-2300 m (5600-7500 ft) |
Distribution |
CO
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AZ; NV; UT |
Discussion | Of conservation concern. Boechera crandallii is a sexual diploid known only from the Gunnison Basin of west-central Colorado. Despite its narrow range, the species appears to hybridize frequently, most notably with B. pallidifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fruiting pedicel length, considered diagnostic by R. C. Rollins (1993) and N. H. Holmgren (2005b), occasionally fails to separate Boechera gracilipes from the closely-related B. fendleri. The two are consistently distinguished by trichome characters. In B. gracilipes, basal leaves usually lack prominent cilia and surfaces are persistently pubescent with at least some 3-rayed trichomes. Also, stems are rather densely pilose proximally, with the largest trichomes more than 0.9 mm. By contrast, basal leaves of B. fendleri always have prominent cilia, surfaces are often glabrescent, and 3-rayed trichomes usually are rare or absent. Stems of the latter are proximally hirsute to hispid, with the largest trichomes less than 0.9 mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 372. | FNA vol. 7, p. 381. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis crandallii, Arabis stenoloba | Arabis gracilipes, Arabis arcuata var. longipes, Arabis perennans var. longipes |
Name authority | (B. L. Robinson) W. A. Weber: Phytologia 51: 369. (1982) | (Greene) Dorn: Brittonia 55: 3. (2003) |
Web links |