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Crandall's rockcress

California rockcress

Habit Perennials; long-lived; (often cespitose); sexual; caudex somewhat woody. Perennials; usually long-lived; apomictic; caudex woody.
Stems

usually 2–5 per caudex branch, arising from margin of rosette near ground surface, (1–)1.5–4 dm, densely pubescent proximally, trichomes short-stalked, 5–8-rayed, 0.1–0.2 mm, moderately to sparsely pubescent distally.

usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, (2–)3.5–13 dm, densely pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.3–0.9 mm, moderately to sparsely pubescent distally.

Basal leaves

blade narrowly oblanceolate, 1.5–3(–5) mm wide, margins entire, ciliate along petiole base, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 5–8-rayed 0.1–0.2 mm.

blade linear to oblanceolate, 3–10(–13) mm wide, margins entire or denticulate, ciliate along petiole, trichomes to 1.3 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.2–0.5 mm.

Cauline leaves

5–14, not concealing stem;

blade auricles 0.1–0.5 mm, surfaces of distalmost leaves pubescent.

12–35(–55), often concealing stem proximally;

blade auricles 1–5(–6) mm, surfaces of distalmost leaves pubescent.

Racemes

10–30-flowered, usually unbranched.

30–120-flowered, usually unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals usually white, 5–7 × 1–2 mm, glabrous;

pollen ellipsoid.

divaricate-ascending at anthesis;

sepals pubescent;

petals usually purple, rarely pinkish, 9–14 × 1.5–3 mm, glabrous or sparsely pubescent (trichomes scattered abaxially);

pollen spheroid.

Fruiting pedicels

ascending to divaricate-ascending, straight, 5–10 mm, pubescent, trichomes branched.

descending to horizontal, usually recurved, rarely straight, 4–20 mm, pubescent, trichomes appressed, 3–7-rayed.

Fruits

ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 3–5.5 cm × 0.9–1.2 mm;

valves glabrous;

ovules 56–84 per ovary;

style 0.1–0.5 mm.

widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (6–)8–12 cm × 1.5–2.5 mm;

valves glabrous or trichomes sometimes scattered;

ovules 140–180 per ovary;

style 0.05–0.3 mm.

Seeds

uniseriate, 0.8–1 × 0.6–0.9 mm;

wing continuous (rarely absent), to 0.1 mm wide.

uniseriate or sub-biseriate, 1.7–2 × 1.2–1.7 mm;

wing continuous, 0.2–0.4 mm wide.

2n

= 14.

Boechera crandallii

Boechera californica

Phenology Flowering May–Jun. Flowering Mar–Jun.
Habitat Rocky slopes and gravelly soil in sagebrush, mountain shrub, open conifer forests Rocky slopes and gravelly soil in desert chaparral and oak woodlands
Elevation 2000-2700 m (6600-8900 ft) 300-2300 m (1000-7500 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Boechera crandallii is a sexual diploid known only from the Gunnison Basin of west-central Colorado. Despite its narrow range, the species appears to hybridize frequently, most notably with B. pallidifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera californica is an apomictic species that arose through hybridization between B. arcuata and B. pulchra (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). In the flora area, B. californica is known only from Los Angeles, Riverside, San Bernardino, and San Diego counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 372. FNA vol. 7, p. 369.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis crandallii, Arabis stenoloba Arabis sparsiflora var. californica
Name authority (B. L. Robinson) W. A. Weber: Phytologia 51: 369. (1982) (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 260. (2007)
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