Asteraceae tribe Heliantheae subtribe Pectidinae
Adenophyllum
dogweed
erect, branched from bases or throughout.
mostly cauline, sometimes basal or basal and cauline; mostly opposite (distal sometimes alternate) or mostly alternate;
petiolate or sessile;
blades usually elliptic, filiform, lanceolate, linear, or ovate, often 1(–2)-pinnately lobed, ultimate margins entire or toothed (often proximally bristly-ciliate), faces glabrous or puberulent to tomentose (usually bearing oil-glands, embedded pellucid glands filled with strong-scented, essential oils).
cauline; mostly opposite (distal sometimes alternate);
petiolate or sessile;
blades (or lobes) ovate, oblanceolate, or linear, 1-pinnately lobed, ultimate margins ± toothed (teeth often bristle-tipped), faces glabrous or ± hirtellous to scabrellous (oil-glands usually borne at bases of lobes and subterminally near apices).
campanulate, cylindric, hemispheric, obconic, or turbinate.
campanulate to obconic [hemispheric], [5–]7–20+ mm diam.
flat or convex to hemispheric or conic, epaleate (receptacles sometimes pitted, pit margins sometimes fimbrillate to laciniate).
convex, ± pitted (socket margins fimbrillate to setose), epaleate.
0 or 1–21+, pistillate, fertile;
corollas yellow to orange (sometimes with brown or mostly brown) or white to pink, magenta, or reddish purple.
usually 7–16[–16], rarely 0, pistillate, fertile;
corollas yellow to orange or red-orange.
1–120+, usually bisexual and fertile [functionally staminate];
corollas yellow to orange or brown, or pinkish to purplish or reddish, tubes much shorter than to much longer than cylindric to funnelform throats, lobes 5, deltate to lance-linear (equal or unequal);
anther thecae pale or ± darkened;
stigmatic papillae in 2 lines.
25–70[100+], bisexual, fertile;
corollas yellow to yellow-orange, tubes shorter than cylindric throats, lobes 5, lance-ovate to lance-linear.
falling or persistent, 3–21+ in 1–2 series (distinct or ± connate, mostly linear to lanceolate, usually subequal, usually some or all bearing pellucid oil-glands as in leaves).
persistent, 8–20+ in ± 2 series (weakly connate 1/3–3/4 their lengths, separating in age, mostly lanceolate to linear, margins of outer distinct to bases or nearly so, oil-glands in faces and near margins).
0 or of 1–22 bractlets.
of 3–5 or 12–22 linear to subulate bractlets (sometimes with 1–2 lobes, often bristle-tipped, usually bearing oil-glands).
radiate or discoid (sometimes radiant, e.g., in Thymophylla), usually borne singly, sometimes in loose to crowded, corymbiform or cymiform arrays.
usually radiate, rarely discoid, borne singly.
mostly clavate to obpyramidal (lengths usually 2–3+ times diams.), glabrous or hairy;
pappi usually persistent, of 20–50 bristles in 1–2+ series, or of 5–20 (sometimes aristate) scales in 1–2 series, or combinations of bristles and scales in 1–2+ series, rarely coroniform or 0.
obpyramidal [obconic], glabrous or ± strigose or sericeous;
pappi persistent, of 8–20 scales in 1–2 series (single scales muticous, or unequally 3-aristate, or comprising 5–11 basally connate bristles).
= 7, 13.
Asteraceae tribe Heliantheae subtribe Pectidinae
Adenophyllum
Genera 22, species 225 (9 genera, 38 species in the flora).
Circumscription of Pectidinae has been fairly constant for decades. The group is sometimes given tribal rank, as Tageteae. Here, we have followed H. Robinson (1981). The group is usually distinguished on the basis of schizogenous glands containing essential oils (mostly terpenes) in tissues of leaves and/or phyllaries (sometimes in tissues of other organs as well).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species 10 (3 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaves opposite (blade margins proximally bristly-ciliate; ray florets borne on bases ofsubtending phyllaries; style branches of bisexual florets knoblike) | Pectis |
1. Leaves opposite or mostly alternate (blades often lobed, margins sometimes bristly-ciliate; ray florets borne on receptacles, not on bases of subtending phyllaries; style branches of bisexual florets linear) | → 2 |
2. Phyllaries distinct to bases or nearly so | → 3 |
2. Phyllaries ± connate 1/3–7/8+ their lengths (margins of outer may be free to bases) | → 6 |
3. Calyculi of (0–)1–9, deltate, lanceolate, or linear bractlets; pappi wholly or partly of scales (individual scales sometimes comprising 5–10, basally connate bristles) | → 4 |
3. Calyculi 0; pappi wholly of distinct bristles | → 5 |
4. Ray corollas yellow to orange | Dyssodia |
4. Ray corollas whitish with pinkish or purplish stripes | Nicolletia |
5. Heads radiate | Chrysactinia |
5. Heads discoid | Porophyllum |
6. Calyculi 0; pappi of 2–5(–10) elements in ± 1 series (usually 0–5+ oblong to lanceolate, erose-truncate to laciniate, plus 0–2+ longer, subulate or aristate, some or all sometimes connate) | Tagetes |
6. Calyculi usually of (1–)5–8(–22) bractlets, rarely 0; pappi usually of 8–20 elements in 2 series, rarely coroniform | → 7 |
7. Bractlets of calyculi subulate or pectinate | Dysodiopsis |
7. Bractlets of calyculi (deltate to subulate) not pectinate | → 8 |
8. Plants (20–)30–70+ cm; involucres (7–18 ×) 5–12 mm; phyllaries weakly connate 1/3–2/3 their lengths | Adenophyllum |
8. Plants mostly (1–)5–30 cm; involucres (4–6 ×) 2–7 mm; phyllaries strongly connate 2/3–7/8 their lengths (margins of outer may be free to bases) | Thymophylla |
1. Annuals; calyculus bractlets eglandular; pappi: outer scales erose, inner each unequally 3-aristate | A. wrightii |
1. Perennials or subshrubs; calyculus bractlets bearing oil-glands; pappi: each scale comprising 5–11 basally connate bristles | → 2 |
2. Leaves 3–5-lobed (lobes linear to cuneate or oblanceolate, ultimate margins entire or toothed); disc florets 25–40; pappi of 8–12 scales | A. porophylloides |
2. Leaves not lobed (blades ovate to oblanceolate, margins toothed); disc florets 50–80; pappi of 15–20 scales | A. cooperi |
CA
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